MTSU ALERT: MTSU will remain closed Friday, March 6, due to weather. Classes are cancelled and offices closed. TSSAA games will proceed. Check for more information.
Department of Biology

Dr. R. Stephen HowardHOWARD

Phone: (615) 898-2044
Fax: (615) 898-5093
Office Location: SCI 1051
MTSU Box X087 Mufreesboro, TN 37132

B.S. Magna Cum Laude (Biology), Eastern Kentucky University
M.S. (Biology), Eastern Kentucky University
Ph.D. (Biology), Indiana University

Host-parasite interactions, evolutionary theory


Eric J. Nordberg, Tom A. Blanchard, Vincent A. Cobb, A. Floyd Scott, and R. Stephen Howard. 2013. Distribution of a Non-Native Gecko (Hemidactylus turcicus) in Tennessee. Tennessee Academy of Science

DA Adams, JL Walck, RS Howard, P Milberg. 2012. Forest Composition and Structure on Glade-forming Limestones in Middle Tennessee.  Castanea 77 (4), 335-347

JM Nunley, MF Owens, RS Howard. 2011. The effects of information and competition on racial discrimination: Evidence from a field experiment.  Journal of Economic Behavior & Organization 80 (3), 670-679

SG Johnson, RS Howard. 2007. Contrasting patterns of synonymous and nonsynonymous sequence evolution in asexual and sexual freshwater snail lineages.  Evolution 61 (11), 2728-2735

R. S. Howard, C. M. Lively. 2004. Good vs complementary genes for parasite resistance and the evolution of mate choice.  BMC evolutionary biology 4 (1), 48

J. F. Fontanari, A. Colato, and R. S. Howard. 2003. Mutation accumulation in growing asexual lineages. Physical Review Letters 91(21) 218101

R. S. Howard and C. M. Lively. 2003. Opposites attract? Mate choice for parasite evasion and the evolutionary stability of sex. Journal of Evolutionary Biology 16:681-689

R. S. Howard and C. M. Lively. 2002. The Ratchet and the Red Queen: the maintenance of sex in parasites. Journal of Evolutionary Biology 15:648-656

R. S. Howard and C. M. Lively. 1998. The maintenance of sex by parasitism and mutation accumulation under epistatic fitness functions. Evolution 52 (2):604-610 {PDF}

R. S. Howard. 1994. Selection against deleterious mutations and the maintenance of biparental sex. Theoretical Population Biology 45:313-323 {PDF}

R. S. Howard and C. M. Lively. 1994. Parasitism, mutation accumulation and the maintenance of sex. Nature. (367) 554-557

C. M. Lively and R. S. Howard. 1994. Selection by parasites for clonal diversity and mixed mating.  Philosophical transactions of the Royal Society of London, Series B. (346) 271-280

European Society for Evolutionary Biology Society for the Study of Evolution American Association for the Advancement of Science

General Ecology (4240/5240) Graduate Seminar (6650) Speciation (6200)

Full member of the Graduate Faculty