DR. R. STEPHEN HOWARD
Professor

CONTACT:
E-mail: Steve.Howard@mtsu.edu
Phone: (615) 898-2044
Fax: (615) 898-5093
Office Location: 
Haynes House #9 
MTSU Box X087
Mufreesboro, TN 37132

EDUCATION:
B.S. Magna Cum Laude (Biology), Eastern Kentucky University
M.S. (Biology), Eastern Kentucky University
Ph.D. (Biology), Indiana University

RESEARCH INTERESTS:
Host-parasite interactions, evolutionary theory

REPRESENTATIVE PUBLICATIONS:
(Visit my web site at www.mtsu.edu/~rshoward for complete list.)

J. F. Fontanari, A. Colato, and R.S. Howard. 2003. Mutation accumulation in growing asexual lineages. Physical Review Letters 91(21) 218101

R. S. Howard and C. M. Lively. 2003. Opposites attract? Mate choice for parasite evasion and the evolutionary stability of sex. Journal of Evolutionary Biology 16:681-689

R. S. Howard and C. M. Lively. 2002. The Ratchet and the Red Queen: the maintenance of sex in parasites. Journal of Evolutionary Biology 15:648-656

R. S. Howard and C. M. Lively. 1998. The maintenance of sex by parasitism and mutation accumulation under epistatic fitness functions. Evolution 52 (2):604-610 {PDF}

R. S. Howard. 1994. Selection against deleterious mutations and the maintenance of biparental sex. Theoretical Population Biology 45:313-323 {PDF}

PROFESSIONAL SOCIETIES:
European Society for Evolutionary Biology
Society for the Study of Evolution
American Association for the Advancement of Science

COURSES TAUGHT:
General Ecology (4240/5240)
Graduate Seminar (6650)
Speciation (6200)

OTHER:
Full member of the Graduate Faculty