Dr. Andrew Van Zandt Brower

Professor

Dr. Andrew Van Zandt Brower
615-898-2064
Room 1053, Science Building (SCI)
MTSU Box 60, Murfreesboro, TN 37132

Degree Information

  • Ph.D., Cornell University, NY (1994)
  • M.E.S., Yale University (1987)
  • B.S., Yale University (1985)

Areas of Expertise

My empirical research is focused on the molecular systematics, biogeography and evolution of butterflies, mainly in the family Nymphalidae. I am also interested in theoretical principles and methods of systematic biology and biological nomenclature.

Biography

Other Active Appointments
Research Associate, Dept. of Entomology, National Museum of Natural History,
Research Associate, Dept. of Invertebrates, American Museum of Natural History,
Associate Editor, Cladistics
Associate Editor , Systematics and Biodiversity
Rhopalocera TWIG leader, NSF ATOL Lepidoptera Grant (Mitter, Regier, Cummings, Parr, Weller, Davis)
Taxonomy Advisor (Rhopalocera), NCBI GenBank

Past Professi...

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Other Active Appointments
Research Associate, Dept. of Entomology, National Museum of Natural History,
Research Associate, Dept. of Invertebrates, American Museum of Natural History,
Associate Editor, Cladistics
Associate Editor , Systematics and Biodiversity
Rhopalocera TWIG leader, NSF ATOL Lepidoptera Grant (Mitter, Regier, Cummings, Parr, Weller, Davis)
Taxonomy Advisor (Rhopalocera), NCBI GenBank

Past Professional Appointments
Rice Chair of Systematic Entomology, Oregon State University
Assistant Professor (Nov. 1997- June 2006)

Curator of Lepidoptera, Oregon State Arthropod Collection, Oregon State University
(Nov. 1997-June 2006 )

Smithsonian Postdoctoral Fellow, National Museum of Natural History (Mar.-Nov. 1997)

Kalbfleisch Fellow, American Museum of Natural History (May 1996-Feb. 1997 )
Postdoctoral Fellow, American Museum of Natural History (Oct. 1995-April 1996)
NSF Postdoctoral Fellow, American Museum of Natural History (Oct. 1993-Sept. 1995)

Education:

Ph.D., 1994, Cornell University, Section of Ecology and Systematics Dissertation: "Phylogeny and biogeography of Heliconius butterflies inferred from mitochondrial DNA sequences." (275 pp.) Graduate committee: R. G. Harrison (major   advisor), A. R. McCune, J. K. Liebherr, C. F. Aquadro

M.E.S., 1987, Yale School of Forestry and Environmental Studies (concentrations in conservation biology and museology) advisors C. L. Remington, S. R. Kellert

B.A., 1985, Yale College (major in biology) advisor C. L. Remington

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Publications

Peer-Reviewed Research Publications (in chronological order):

PDFs for many of these are available at ResearchGate.

Brower, AVZ and TM Boyce. 1991. Mitochondrial DNA variation in monarch butterflies. Evolution 45:1281-1286.

Brower, AVZ. 1994. Rapid morphological radiation and convergence in geographical races of the butterfly, Heliconiu...

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Peer-Reviewed Research Publications (in chronological order):

PDFs for many of these are available at ResearchGate.

Brower, AVZ and TM Boyce. 1991. Mitochondrial DNA variation in monarch butterflies. Evolution 45:1281-1286.

Brower, AVZ. 1994. Rapid morphological radiation and convergence in geographical races of the butterfly, Heliconius erato, inferred from patterns of mitochondrial DNA evolution. Proc. Natl. Acad. Sci. USA 91:6491-6495.

Brower, AVZ. 1994. Phylogeny of Heliconius butterflies inferred from mitochondrial DNA sequences. Mol. Phylogenet. Evol. 3:159-174.

Brower, AVZ. 1994. The case of the missing H: Heliconius charithonia (L., 1767), not "Heliconius charitonia (L. 1767)". J. Lepidopterists' Soc. 48:166-168.

Brower, AVZ. and R DeSalle. 1994. Practical and theoretical considerations for choice of a DNA sequence region in insect molecular systematics, with a short review of published studies using nuclear gene regions. Ann. Entomol. Soc. Amer. 87:702-716.

Brower, AVZ. 1995. Locomotor mimicry in butterflies? A critical review of the evidence. Phil. Trans. Royal Soc. Lond. B 347:413-425.

Brower, LP, LS Fink, AVZ Brower, K Leong, K Oberhauser, S Altizer, O Taylor, D Vickerman, WH Calvert, T Van Hook, A Alonso M, SB Malcolm, DF Owen and MP Zalucki. 1995. On the dangers of interpopulational transfers of monarch butterflies.BioScience 45:540-544.

Brower, AVZ. 1996. Parallel race formation and the evolution of mimicry in Heliconius butterflies: a phylogenetic hypothesis from mitochondrial DNA sequences. Evolution 50:195-221.

Brower, AVZ. 1996. A new mimetic species of Heliconius (Lepidoptera: Nymphalidae), from southeastern Colombia, revealed by cladistic analysis of mitochondrial DNA sequences. Zool. J. Linn. Soc. 116:317-332.

Brower, AVZ, R DeSalle and AP Vogler. 1996. Gene trees, species trees, and systematics: a cladistic perspective. Annu. Rev. Ecol. Syst. 27:423-450.

Brower, AVZ and VA Schawaroch. 1996. Three steps of homology assessment. Cladistics 12:265-272.

Miller, JS, AVZ Brower and R DeSalle. 1997. Phylogeny of the neotropical moth tribe Josiini (Notodontidae: Dioptinae): Comparing and combining evidence from DNA sequences and morphology. Biol. J. Linn. Soc. 60:297-316.

Brower, AVZ. 1997. The evolution of ecologically important characters in Heliconius butterflies (Lepidoptera: Nymphalidae): a cladistic review. Zool. J. Linn. Soc. 119:457-472.

Brower, AVZ and MG Egan. 1997. Cladistics of Heliconius butterflies and relatives (Nymphalidae: Heliconiiti): the phylogenetic position of Eueides based on sequences from mtDNA and a nuclear gene. Proc. Royal Soc. Lond. B 264:969-977.

DeSalle, R, AVZ Brower, R Baker and J Remsen. 1997. A hierarchical view of the Hawaiian Drosophilidae. Pacific Science 51:462-474 (invited paper from the symposium, “The legacy of RCL Perkins: 100 years of Hawaiian entomology,” XX Int. Congr. Entomol., Florence, Italy, August 1996).

DeSalle, R and AVZ Brower. 1997. Process partitions, congruence and the independence of characters: inferring relationships among closely-related Hawaiian Drosophila from multiple gene regions. Syst. Biol. 46:751-764.

Brower, AVZ and R DeSalle. 1998. Patterns of mitochondrial vs. nuclear DNA sequence divergence among nymphalid butterflies: the utility of Wingless as a source of characters for phylogenetic inference. Insect Mol. Biol.7:1-10.

Brower, AVZ and KR Sime. 1998. A reconsideration of mimicry and aposematism in caterpillars of the Papilio machaon group. J. Lepidopterists’ Soc. 52:206-212.

Sime, KR and AVZ Brower 1998. Explaining the latitudinal gradient anomaly in ichneumonid species richness: evidence from butterflies. J. Animal Ecology 67:387-399.

Brower, AVZ. 1998. How NOT to use systematics to test adaptive hypotheses: comments on Wink and Nickisch-Rosenegk (1997). J. Chemical Ecology 24:1279-1284.

Brower, AVZ. 1999. Delimitation of phylogenetic species with DNA sequences: a critique of Davis and Nixon’s population aggregation analysis. Syst. Biol. 48:199-213.

Brower, AVZ. 2000. Evolution is not a necessary assumption of cladistics. Cladistics 16:143-154.

Brower, AVZ. 2000. Homology, and the inference of systematic relationships: some historical and philosophical perspectives. Pp. 10-21 in RW Scotland and RT Pennington (eds.), Homology and Systematics: Coding characters for phylogenetic analysis. Chapman & Hall.

Campbell, DL, AVZ Brower and NE Pierce. 2000. Molecular evolution of the Wingless gene and its implications for the phylogenetic placement of the butterfly family Riodinidae (Lepidoptera: Papilionoidea). Mol. Biol. Evol.17:684-696.

Brower, AVZ. 2000. On the validity of Heliconius tristero Brower and Heliconius melpomene mocoa Brower, with a discussion of the species concept in Heliconius Kluk (Lepidoptera: Nymphalidae). Proc. Ent. Soc. Wash. 102:678-687.

Brower, AVZ. 2000. Phylogenetic relationships among the Nymphalidae (Lepidoptera), inferred from partial sequences of the wingless gene. Proc. R. Soc. Lond. B 267:1201-1211.

Brower, AVZ. 2002. Cladistics, populations and species in geographical space: the case of Heliconius butterflies. Pp.5-16 in R DeSalle, WC Wheeler and G Giribet (eds.), Molecular Systematics and Evolution: theory and practice. Birkhaüser Verlag.

Goldstein, PZ and AVZ Brower. 2002. Molecular systematics and the origin of species: new syntheses or methodological introgressions? Pp. 147-161 in R DeSalle, WC Wheeler and G Giribet (eds.), Molecular Systematics and Evolution: theory and practice. Birkhaüser Verlag.

Brower, AVZ 2002. Cladistics, phylogeny, evidence and explanation: a reply to Lee. Zoologica Scripta 31:221-223.

Brower, AVZ and M Jeansonne. 2004. Geographical populations and “subspecies” of New World monarch butterflies (Danaus plexippus (L.) Nymphalidae) share a recent origin and are not phylogenetically distinct. Ann. Entomol. Soc. Amer. 97: 519-523.

Brower, AVZ. 2004. Comment on “Molecular phylogenies link rates of evolution and speciation.” Science 303:173c. www.sciencemag.org/cgi/content/full/303/5655/173c

Leathers, JW, DD Judd, and AVZ Brower. 2005. A review of the species of the New World braconid genus Cyclaulacidea (Hymenoptera) with key and descriptions of nine new species. J. Hymenoptera Research 14:151-176.

Silva-Brandão, KL, AVL Freitas, AVZ Brower and VN Solferini. 2005. Phylogenetic relationships of the New World Troidini swallowtails (Lepidoptera: Papilionidae) based on COI, COII, and EF1a genes. Mol. Phylogenet. Evol. 36:468-483.

Wahlberg, N., MF Braby, AVZ Brower, R de Jong, M-M Lee, S Nylin, NE Pierce, FAH Sperling, R Vila, AD Warren and E Zakharov. 2005. Synergistic effects of combining morphological and molecular data in resolving the phylogeny of butterflies and skippers. Proc. R. Soc. Lond. B 272:1577-1586.

Wahlberg, N, AVZ Brower and S Nylin. 2005. Phylogenetic relationships of tribes and genera in the subfamily Nymphalinae ((Lepidoptera: Nymphalidae) based on three gene sequences. Biol. J. Linn. Soc. 86:227-251.

Whinnett A, AVZ Brower, M-M Lee, KR Willmott, and J Mallet 2005. The phylogenetic utility of Tektin, a novel region for inferring systematic relationships amongst Lepidoptera. Ann. Entomol. Soc. Amer. 98:873-886.

Whinnett A, KR Willmott, AVZ Brower, F Simpson, M Zimmermann, G Lamas, and J Mallet. 2005. Mitochondrial DNA provides an insight into the mechanisms driving diversification in the ithomiine butterfly Hyposcada anchiala (Lepidoptera: Nymphalidae, Ithomiinae). European J. Entomol. 102:633-639.

Brower, AVZ. 2006. The how and why of branch support and partitioned branch support, with a new index to assess partition incongruence. Cladistics 22:378-386.

Brower, AVZ. 2006. Problems with DNA barcodes for species delimitation: “ten species” of Astraptes fulgerator reassessed (Lepidoptera: Hesperiidae). Systematics and Biodiversity 4:127-132.

Brower, AVZ, AVL Freitas, M-M Lee, KL Silva Brandão, A Whinnett and KR Willmott 2006. Phylogenetic relationships among the Ithomiini (Lepidoptera: Nymphalidae) inferred from one mitochondrial and two nuclear gene regions. Systematic Entomology 31:288-301.

Peña, C, N Wahlberg, E Weingartner. U Kodandaramaiah, S Nylin, AVL Freitas and AVZ Brower. 2006. Higher level phylogeny of Satyrinae butterflies (Lepidoptera: Nymphalidae) based on DNA sequence data. Mol. Phylogenet. Evol. 40:29-49.

Simonsen, TJ, N Wahlberg, AVZ Brower and R de Jong 2006. Molecules, morphology and fritillaries: a combined approach towards the phylogeny of Argynnini (Lepidoptera: Nymphalidae). Insect Systematics and Evolution 37:405-418.

Beltrán, M, CD Jiggins, AVZ Brower, E Bermingham and J Mallet. 2007. Do pollen feeding, pupal-mating and larval gregariousness have a single origin in Heliconius butterflies? Inferences from multilocus DNA sequence data. Biol. J. Linn. Soc. 92:221-239.

Elias, M, RI Hill, KR Willmott, KK Dasmahapatra, AVZ Brower, J Mallet and CD Jiggins. 2007. Limited performance of DNA barcoding in a diverse community of tropical butterflies. Proc. Royal Soc. London B 274:2881-2889 (on line: doi:10.1098/rspb2007.1035)

Frentiu, FD, GD Bernard, MP Sison-Mangus, AVZ Brower and AD Briscoe. 2007. Gene duplication is an evolutionary mechanism for expanding spectral diversity in the long wavelength photopigments of butterflies. Mol. Biol. Evol. 24:2016-2028.

Warren, AD, J Ogawa and AVZ Brower. 2008. Phylogenetic relationships of subfamilies and circumscription of tribes in the family Hesperiidae (Lepidoptera: Hesperioidea). Cladistics 24:642-676.

Townzen, JS, AVZ Brower and DD Judd. 2008. Identification of mosquito bloodmeals using mitochondrial cytochrome oxidase subunit I and cytochrome b gene sequences. Med. Vet. Entomol. 22:386-393.

Elias, M., M Joron, KR Willmott, V Kaiser, KL Silva-Brandão, C Arias, LM Gomez-Piñeres, S Uribe, AVZ Brower, AVL Freitas and CD Jiggins. 2009. Out of the Andes: patterns of diversification in clearwing butterflies. Mol. Ecol. 18:1716-1729.

Warren, AD, J Ogawa and AVZ Brower 2009. Revised classification of the family Hesperiidae (Lepidoptera: Hesperioidea) based on combined molecular and morphological data. Systematic Entomology 34:467-523.

Brower, AVZ. 2009. Heliconius tristero Brower, 1996 and Heliconius melpomene mocoa Brower, 1996 (Lepidoptera: nymphalidae): proposed conservation by suppression of Heliconius melpomene bellula Brown, 1979. Bull. Zoological Nomenclature 66:256-260.

Wahlberg, N, J Leneveu, U Kodandaramaiah, C Peña, S Nylin, ALV Freitas and AVZ Brower. 2009. Nymphalid butterflies diversify following near demise at the Cretaceous/Tertiary boundary. Proc. Royal Soc. London B 276:4295-4302.

Brower, AVZ. 2010 Stability, replication, pseudoreplication and support. Cladistics 25:112-113. Brower, AVZ, N Wahlberg, JR Ogawa, M Boppré and RI Vane-Wright.. 2010. Phylogenetic relationships among genera of danaine butterflies (Lepidoptera: Nymphalidae) as implied by morphology and DNA sequences. Systematics and Biodiversity 8:75-89.

Brower, AVZ. 2010. Alleviating the taxonomic impediment of DNA barcoding: names for ten species of “Astraptes fulgerator” (Lepidoptera: Hesperiidae: Eudaminae). Systematics and Biodiversity 8:485-491.

Brower, AVZ. 2011. Hybrid speciation in Heliconius butterflies: a review and critique of the evidence. Genetica 139:589-609; online version DOI: 10.1007/s10709-010-9530-4

Rindal, E and AVZ Brower. 2011. Do model-based phylogenetic analyses outperform parsimony? A test with empirical data. Cladistics 27:331-334; online version DOI: 10.1111/j.1096-0031.2010.00342.x

Brower, AVZ. 2011. Repeatability and reality. Cladistics 27:447-448; online version DOI: 10.1111/j.1096-0031.2011.00354.x

Brower, AVZ and RI Vane-Wright. 2011. Anomalous areas and awkward ages: concerns about over-reliance on model based biogeographical and temporal inferences. Systematic Entomology 36:601-603

Brower, AVZ. 2012. The meaning of “phenetic”. Cladistics 28:113-114, online version DOI: 10.1111/j.1096-0031.2011.00374.x.

Brower, AVZ and MCC de Pinna. 2012. Homology and errors. Cladistics 28:529-538, online version DOI: 10.1111/j.1096-0031.2012.00398.x

Brower, AVZ 2013. Introgression of wing pattern alleles and speciation via homoploid hybridization in Heliconius butterflies – a review of evidence from the genome. Proc. Royal Soc. London B 280: 20122302

Brower, AVZ and E Rindal. 2013. Reality check: a reply to Smith. Cladistics 29:464-465, DOI: 10.1111/cla.12016

Brower, AVZ and MCC de Pinna. 2014. About nothing. Cladistics 30: 330-336.

Brower, AVZ, KR Willmott, KL Silva-Brandão, IJ Garzón-Orduña, AVL Freitas 2014. Phylogenetic relationships of ithomiine butterflies (Lepidoptera: Nymphalidae: Danainae) as implied by combined morphological and molecular data. Systematics and Biodiversity 12: 133-147; 259-260 (corrigenda). DOI 10.1080/14772000.2014.899650

Garzón-Orduña, IJ, JE Benetti-Longhini and AVZ Brower. 2014. Timing the diversification of the Amazonian biota: butterfly divergences are consistent with Pleistocene refugia. J. Biogeography 41: 1631-1638, DOI: 10.1111/jbi.12330

Brower, AVZ. 2014 Taxic and transformational homology revisited. Cladistics 31: 197-201, DOI: 10.1111/cla.12076

Garzón-Orduña, IJ, JE Benetti-Longhini and AVZ Brower. 2015. Competing paradigms of Amazonian diversification and the Pleistocene refugium hypothesis. J. Biogeography 42: 1357-1360, DOI: 10.111/jbl.12539.

Garzón-Orduña, IJ, KL Silva-Brandão, KR Willmott, AVL Freitas and AVZ Brower. 2015. An alternative, plant-based time-tree implies conflicting dates for the diversification of ithomiine butterflies (Lepidoptera: Nymphalidae: Danainae). Systematic Biology 64: 752-767, DOI:10.1093/sysbio/syv032.

Brower, AVZ. 2016. What is a cladogram and what is not? Cladistics 32: 573-576, DOI: 10.1111/cla.12144

Brower, AVZ. 2016. Rethinking tree-thinking: cladograms, ancestors and evidence. American Biology Teacher 78: 380-384.

Matz, J, and AVZ Brower. 2016. The South Temperate Pronophilina (Lepidoptera: Nymphalidae: Satyrinae): a phylogenetic hypothesis, redescriptions and revisionary notes. Zootaxa 4125: 1-108.

Brower, AVZ. 2016. Are we all cladists? Pp. 88-114 in: Williams, DM, Wheeler, QD and Schmitt M. (eds.) The Future of Phylogenetic Systematics: the Legacy of Willi Hennig, Cambridge University Press.

Sahoo, RK, AD Warren, N Wahlberg, AVZ Brower, V Lukhtanov and U Kodandaramaiah. 2016. Ten genes and two topologies: an exploration of higher relationships in skipper butterflies (Hesperiidae). PeerJ 4:e2653 https://doi.org/10.7717/peerj.2653

Brower, AVZ and IJ Garzón-Orduña. In press. Missing data, clade support and "reticulation": the molecular systematics of Heliconius and related genera (Lepidoptera: Nymphalidae) reexamined. Cladistics

 

Book:

Schuh, RT and AVZ Brower. 2009. Biological Systematics: principles and applications. 2nd edn. Ithaca, NY: Cornell University Press. 311 pp.  

 

Scientific Correspondence, Published Abstracts, Book Reviews and Nontechnical Articles:

Brower, AVZ. 1986. Update from Moncon II. Atala 14:12-14.

Brower, AVZ. 1993. (Book Review: The Development and Evolution of Butterfly Wing Patterns, by H. F. Nijhout, 1991, Smithsonian Institution Press, Washington). J. N. Y. Entomol. Soc. 101:141-142.

Brower, AVZ. 1994. (Book Review: Biology and Conservation of the Monarch Butterfly, by SB Malcolm and MP Zalucki (eds.), 1992. Los Angeles County Museum Science Series No. 38). J. N. Y. Entomol. Soc. 102:397-399.

Brower, AVZ. 1995. Taxonomic minimalism (A response to Beattie and Oliver 1994). Trends in Ecol. and Evol. 10:203.

Brower, AVZ. 1995. (Book review: Butterflies and Climate Change, by RLH Dennis, 1993. Manchester University Press, Manchester U. K.) J. N. Y. Entomol. Soc. 103:123-125.

Brower, AVZ. 1995. (Book review: The Butterflies of the West Indies and South Florida, by DS Smith et al., 1994, Oxford University Press, Oxford.) J. N. Y. Entomol. Soc. 103:126-128.

Brower, AVZ. 1996. Combining data in phylogenetic analysis (a response to Huelsenbeck et al. 1996). Trends in Ecol. and Evol. 11:334-335.

Brower, AVZ. 1996. (Book review: Molecular Model Systems in the Lepidoptera, by MR Goldsmith and AS Wilkins (eds.), 1995, Cambridge University Press, Cambridge.) Bull. Entomol. Research 86:630-631.

Brower, AVZ. 1996. Butterfly ranching in New Guinea. Faces Magazine October 1996:14.

Brower, LP, LS Fink, AVZ Brower, K Leong, K Oberhauser, S Altizer, O Taylor, D Vickerman, WH Calvert, T Van Hook, A Alonso M, SB Malcolm, DF Owen and MP Zalucki. 1996. Lincoln P. Brower et al. reply (to “Monarch transfer: a real concern?” by JB Keiper). BioScience 46:562-563.

Brower, AVZ. 1997. (Book review: The Butterflies of Venezuela, by AFE Neild, Meridian Publications, London). J. N. Y. Entomol. Soc. 104:236-238.

Brower, AVZ, G Chavarría and DD Judd. 1997. Phylogenetic analysis (response to Hillis, 1997). Science 276: 318.

Brower, AVZ. 1998. (letter to the editor). News Lepid. Soc. 40:10.

Brower, AVZ. 1998. (Book review: Las Mariposas de Chile/The Butterflies of Chile, by L. E. Peña G. and A. J. Ugarte P., Editorial Universitaria, Santiago). J. N. Y. Entomol. Soc. 106:115-116.

Brower, AVZ and DD Judd. 1998. Net-wielding anachronisms? (response to Bull and Wichman, 1998). Science 282:1048.

Brower, AVZ. 1998. A fair book review? A reply to Mr. Wilmott. News Lepid. Soc. 40:106.

Brower, AVZ. 1998. (Book review: Heliconius and Related Genera, by H and R Holzinger, 1994, Sciences Nat, Vennete, France.) J. Lepidopterists’ Soc. 52:120-122.

Brower, AVZ. 1999. (letter to the editor). News Lepid. Soc. 41:93.

Brower, AVZ. 1999 (2000). (Book review: Butterflies of Papua New Guinea, by M. L. Parsons, 1998, Academic Press, San Diego). J. N. Y. Entomol. Soc. 107:410-414.

Brower, AVZ. 2000. To Collaborate, compete, or ignore? International Congress of Entomology XXI Abstracts 2:920

Brower, AVZ. 2000. Phylogeny of Nymphalidae, as inferred from wingless and mtDNA sequences. International Congress of Entomology XXI Abstracts 2:934.

Brower, AVZ. 2001. (Book review: Hawkmoths of the World, by I.J. Kitching and J.-M. Cadiou, 2000, Cornell University Press, Ithaca, NY) J. Lepidopterists’ Soc. 54:101.

Brower, AVZ. 2002. Cladistics with and without metaphysical bombast. P. 219 in Judd, DD and AVZ Brower (eds.). Abstracts of the 20th annual meeting of the Willi Hennig Society. Cladistics 18: 218-236.

Judd, DD and AVZ Brower. 2002. Abstracts of the 20th annual meeting of the Willi Hennig Society. Cladistics 18:218-236.

Brower, AVZ. 2002. Molecular systematics of butterflies: a review and prospectus. 4th International Conference on the Biology of Butterflies Abstracts:11.

Wahlberg, N., AVZ. Brower and S Nylin. 2002. Molecular phylogeny of the subfamily Nymphalinae based on mitochondrial and nuclear DNA sequences. 4th International Conference on the Biology of Butterflies Abstracts: 34.

Brower, AVZ. 2002. The scientific character of evolutionary theory is supported by cladistics, and not the other way around. VI International Congress of Systematics and Evolutionary Biology Abstracts:32.

Brower, AVZ. 2002. Waiting for GloBIS: progress and prospects for web-based butterfly taxonomic resources. VI International Congress of Systematics and Evolutionary Biology Abstracts:57.

Brower, AVZ. 2002. Practical and philosophical benefits of the cladistic (MP) approach for analysis of data from multiple sources. VI International Congress of Systematics and Evolutionary Biology Abstracts:215.

Wahlberg, N, M Braby, A Brower, D Campbell, S Nylin, N Pierce, F Sperling, R Vila, A Warren, W Watt, T Yamauchi, O Yata and E Zakharov. 2002 Towards a better understanding of butterfly phylogeny: the first results of a broad collaboration in the molecular systematics of Papilionoidea and Hesperioidea. VI International Congress of Systematics and Evolutionary Biology Abstracts:58.

Brower, AVZ. 2003. Oregon State Arthropod Collection receives Shepard Collection. News of the Lepid. Soc. 45(4):122.

Brower, AVZ. 2004 Phylogenetic relationships of Ithomiinae and Danainae (Lepidoptera: Nymphalidae) inferred from DNA sequences. P. 588 in Grandcolas, P., Abstracts of the 23rd annual meeting of the Willi Hennig Society. Cladistics 20:583-608.

Brower, AVZ. 2005. Roland Trimen’s observations on toughness in unpalatable African butterflies: a historical annotation to DeVries 2003. Journal of the Lepidopterists’ Society 59:44.

Brower, AVZ. 2005. Molecular systematics of Ithomiini and Danaini (Nymphalidae: Danainae). I Encontro sobre Lepidoptera Neotropicais Abstracts: 28.

Brower, AVZ. 2005. Molecular systematics of butterflies. Lepidopterists’ Society 56th Annual Meeting Abstracts:18.

Brower, AVZ and J Gatesy. 2005. The Partition Congruence Index: a measurement of congruence/conflict among data partitions in simultaneous analysis. Abstracts of the24th Willi Hennig Society Meeting:22-23.

Miller, KL, R Wagner, AVZ Brower, and DD Judd. 2005. Relationships of the nematoceran flies: tanyderids + psychodids, a discarded hypothesis is revived by molecular data. Abstracts of the 24th Willi Hennig Society Meeting:27.

Peña, C., N. Wahlberg, E. Weingartner, U. Kodandaramaiah, S. Nylin, AVL Freitas and AVZ Brower. 2005. Filogenia de mariposas Satyrinae (Lepidoptera: Nymphalidae) basada en sequencias de DNA: un estudio preliminar. I Encontro sobre Lepidoptera Neotropicais Abstracts 26.

Brower, AVZ. 2006. (Book review: The genus Adelpha: its systematics, biology and biogeography, by KR Willmott, 2003, Scientific Publishers, Gainesville, FLA). Ann. Entomol. Soc. Amer. 99:184-185.

Brower, AVZ. 2006. Comment on Tree Thinking,. Electronic letter posted by Science, http://www.sciencemag.org/cgi/eletters/310/5750/979#3414 (published online 22 March 2006).

Brower LP, K Oberhauser, M Boppré, AVZ Brower, and RI Vane-Wright. 2007. Monarch sex: ancient rites, or recent wrongs? Antenna 31:12-18.

Elias, M, M Joron, KR Willmott, V Kaiser, KL Silva-Brandão, AVL Freitas, C Arias Mejía, LM Gomez Pineres, AVZ Brower. & C Jiggins. 2007 Phylogenetic hypothesis, pattern of speciation and evolution of wing pattern in neotropical Napeogenes butterflies (Lepidoptera: Nymphalidae). In 7th International Workshop on the Molecular Biology and Genetics of the Lepidoptera August 20–26, 2006, Orthodox Academy of Crete, Kolympari, Crete, Greece. 52pp., vol. 7:29 (ed. K. Iatrou & P. Couble), pp. 13-14: Journal of Insect Science.

Brower, AVZ. 2007. Tree of Life for butterflies makes butterfly relationships accessible at the click of a mouse. News Lepid. Soc. 49:122.

Brower, AVZ. 2009. Science as a pattern (book review of Foundations of Systematics and Biogeography, by DM Williams and MC Ebach, 2008. Springer, NY). Systematics and Biodiversity 7: 345-346.

Brower, AVZ. 2010. Should evolutionary theory evolve? The Scientist 24:14.

Brower, AVZ. 2011. Really learning biology. The Scientist 25:12

Brower, AVZ. 2012. A picture is worth a thousand words (book review of Trees of Life: a Visual History of Evolution, by TW Pietsch, 2012, John Hopkins, Baltimore, MD). Cladistics 28: 655-657.

Brower, AVZ. 2013. Communing with the ancestors (book review of Tree Thinking: an Introduction to Phylogenetic Biology, by DA Baum and SD Smith, 2012, Roberts & Co., Greenwood Village, CO) Cladistics 30: 107-11, DOI: 10.1111/cla/12033

Brower, AVZ. 2014. Willi Hennig at 100 (book review of From Taxonomy to Phylogenetics – Life and Work of Willi Hennig, by Michael Schmitt. Koninklijke Brill NV, Leiden, Netherlands) Cladistics 30: 224-225.

Brower, AVZ. 2014. “A metaphysical research programme” (book review of Phylogenetics: Theory and Practice of Phylogenetic Systematics, by EO Wiley and BS Lieberman. Wiley-Blackwell, London) Cladistics 31: 226-230, DOI: 10.1111/cla.12072.

Brower, AVZ. 2014 “Theory free”? (book review of The Nature of Classification, by JS Wilkins and MC Ebach. Palgrave-MacMillan, New York) Cladistics 31: 337-339, DOI: 10.1111/cla.12086

Brower, AVZ. 2015. Paraphylophily (book review of A Framework for Post-Phylogenetic Systematics, by RH Zander. Zetetic Publications, St Louis, MO) Cladistics 31: 575-578, DOI: 10.1111/cla.12096

"the editors" 2016. Editorial. Cladistics 32: 1, DOI: 10.1111/cla.12148

Brower, AVZ. 2016. Trees and more trees (book reviews of Aristotle’s Ladder, Darwin’s Tree, by JD Archibald. Columbia University Press, New York, and The Book of Trees, by M Lima. Princeton Architectural Press, New York). Cladistics 32: 215-218, DOI: 10.1111/cla.12122

Brower, AVZ. 2016. Emergent properties. (book review of Homology, Genes, and Evolutionary Innovation, by GP Wagner. Princeton University Press, Princeton, NJ). Cladistics 32: 577-579, DOI: 10.1111/cla.12152

Brower, AVZ. 2016. (Book review of Butterflies, a Complete Guide to their Biology and Behavior, by D Vane-Wright. Cornell University Press, Ithaca, NY). Quarterly Review of Biology 91:105.

Brower, AVZ. 2016. (Book review of African Queens and Their Kin: a Darwinian Odyssey, by DAS Smith. Brambleby Books, Harpenden, Herts, UK). News Lepid. Soc. 58: 86-87.

Brower, AVZ. 2016. Tree-thinking (Letter in response to "On being a fish" by B Weinstein). Inference 2 (4) http://inference-review.com/article/tree-thinking

Brower, AVZ. 2017. "Parsimony be damned!" (Book review of Ockham's Razors: A User's Guide, by E Sober. Cambridge University Press) Cladistics 2017. Early view: DOI:10.1111/cla.12190  

 

Web Publications:

ToLWeb pages: Brower, AVZ. 2006-2013. Nymphalidae, Pieridae, Hesperiidae, Riodinidae, Lycaenidae, Dioptinae (Notodontidae) (and many included taxa: 2450+ pages) Sept. 2006 – 2013, in The Tree of Life Web Project, http://tolweb.org/

 

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Presentations

Seminars and Invited Symposia:

Symposium on Biogeography, Willi Hennig Society meeting, Buenos Aires, Oct. 2016 (coauthor, talk presented by C Szumik)

Symposium, 7th International Conference on the Biology of Butterflies, Turku, Finland, Aug. 2014 (coauthor, talk presented by IJ Garzón-Orduña)

Plenary talk, Dimensions of Amazonian Biodiversity Symposium, Manaus, Brazil, May 2014 (coauthor, talk presented by IJ Garzón-Orduña)...

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Seminars and Invited Symposia:

Symposium on Biogeography, Willi Hennig Society meeting, Buenos Aires, Oct. 2016 (coauthor, talk presented by C Szumik)

Symposium, 7th International Conference on the Biology of Butterflies, Turku, Finland, Aug. 2014 (coauthor, talk presented by IJ Garzón-Orduña)

Plenary talk, Dimensions of Amazonian Biodiversity Symposium, Manaus, Brazil, May 2014 (coauthor, talk presented by IJ Garzón-Orduña) I

nvited speaker, Willi Hennig Centennial Symposium, Linnean Society, London, Nov. 2013

Plenary speaker, Dimensions of Amazonian Biodiversity Symposium, São Paulo, Brazil, March 2013

Seminar, Ecology and Evolution brown bag, Dept. Biological Sciences, Vanderbilt U., Feb. 2012

Seminar, University of the South, Sewanee TN, Oct. 2011

Invited symposium, Hennig Society meeting, São José do Rio Preto, Brazil, Aug. 2011 (“History and Philosophy of Cladistics”)

Seminar, Dept. Biology, MTSU, Oct. 2010

Invited symposium in honor of Rick Harrison, Cornell University, July 2010

Seminar, Dept. Biology, MTSU, Feb. 2010

Seminar, Dept. Biology, MTSU, Oct. 2009

2 seminars. Dept. Biology, MTSU, Feb. 2009

2 symposia, V International Butterfly Ecology and Evolution Symposium, Rome, July 2007 (coauthor: was unable to attend and present two talks)

Symposium, Naturalis, Leiden, Netherlands, April 2007

Seminar, Naturalis/University of Leiden, Leiden, Netherlands, Feb. 2006

Seminar, Middle Tennessee State University, Murfreesboro TN, Feb. 2006

Invited talk, Entomological Collections Network, Ft. Lauderdale, FL, Nov. 2005

Invited symposium, Entomological Society of America, Ft. Lauderdale, FL, Nov. 2005

Plenary address, Lepidopterists’ Society, Sierra Vista AZ, August 2005

Symposium, “Philosophy and methodology” Willi Hennig Society, Fagernes, Norway, July 2005

Seminar, Naturhistoriska Riksmuseet, Stockholm, Sweden, June 2005

Invited symposium, I Encontro sobre Lepidoptera Neotropicais, Universidade Estadual de Campinas, Brazil, April 2005 (author and coauthor)

Seminar, University of Alaska Museum of the North, Fairbanks, AK, March 2005

Invited symposium, “Lepidoptera Erotica: a passion for butterflies,” Entomological Society of America annual meeting, Salt Lake City, Nov. 2004

Invited symposium, “Classifying things,” History of Science Department, Oregon State U., Oct. 2004

Symposium, XXIV International Congress of Entomology, Brisbane, Australia, August 2004 (symposium title: “Mimicry in the new millennium,” co-organized with Rebecca Simmons)

Symposium, “Phylogenetics and evolution of arthropods,” Willi Hennig Society, Paris, July 2004

Seminar, Universidade Estadual de Campinas, Brazil, March 2004 Invited symposium, Lepidopterists’ Society, Olds Alberta, July 2003 (symposium topic: “The future of Lepidopterology”)

Invited presentation, GART/GloBIS workshop, Stuttgart, Germany, May 2003 Invited

Symposium, International Congress of Systematics and Evolutionary Biology, Patras, Greece, Sept. 2002 (symposium title: “Philosophical Issues in Phylogenetic Inference”)

Invited Symposium, International Congress of Systematics and Evolutionary Biology, Patras, Greece, Sept. 2002 (symposium title, “Incongruence, Data Partitions and Phylogenetic Signal ”)

Invited Symposium, International Congress of Systematics and Evolutionary Biology, Patras, Greece, Sept. 2002 (symposium title: “Butterfly Biodiversity: patterns and processes from Lepidoptera genomics to ecosystems”) (author of one presentation and coauthor of a second)

Plenary Speaker, IV International Butterfly Ecology and Evolution Symposium, Leiden, Netherlands, March 2002

Symposium, “Cladistics and Philosophy” XX Hennig Society meeting, Corvallis, Aug. 2001

Seminar and Lecture, University of Minnesota, Dept. of Entomology, March 2001

Symposium, “The evidence for evolution,” OSU Hashimoto Award Symposium, Nov. 2000

Seminar, OSU Dept. of Zoology, Oct. 2000

Symposium, XXIII International Congress of Entomology, Foz de Iguassu, Brazil, Aug. 2000 (symposium title: “Congruence among data from different sources in systematic entomology,” co-organizer with D. Judd)

Symposium, XXIII International Congress of Entomology, Foz de Iguassu, Brazil, Aug. 2000 (symposium title: “Systematics and phylogeny of Lepidoptera”)

Symposium, Association for Tropical Biology, Bloomington, IN, June 2000

Seminar, OSU Dept. of Entomology, April 2000

Invited talk, OSU Center for Gene Research and Biotech. retreat, Newport OR, Oct. 1999

Invited talk, Lepidopterists’ Society Meeting, Sierra Vista AZ, August 1999 (Symposium topic: species concepts in Lepidoptera)

Seminar, University of Florida, Dept. of Entomology, Nov. 1998

Seminar, University of Arizona Dept. of Entomology, August 1998

Symposium, systematics session, III International Symposium on Butterfly Ecology and Evolution, Crested Butte, CO August 1998

Invited Symposium, Systematics Association, Oxford, U. K. August 1997 (Symposium title: “Character conceptualisation, primary homology and systematic solutions”)

Symposium, Lepidopterists’ Society 50th Anniversary Meeting, New Haven, CT July 1997 (Symposium topic: molecular and morphological systematics of Lepidoptera)

Seminar, University of California ESPM (Entomology), Berkeley, CA April 1997

Seminar, Oregon State University Dept. of Entomology, Corvallis, OR April 1997

Seminar, Iowa State University Dept. of Entomology, Ames, IA April 1997

Seminar, University of Rochester Dept. of Biology, Rochester, NY Feb. 1997

Seminar, Queens College Dept. of Biology, Flushing, NY Feb. 1997

Seminar, American Museum Systematics Discussion Group, NY, NY Sept. 1996

Invited talk, Eastern Great Lakes Molecular Evolution Meeting, Ithaca, NY May 1996

Seminar, Harvard University, Museum of Comparative Zoology, May 1996

Seminar, Columbia University, CERC, March 1996

Seminar, Cornell University, Section of Ecology and Systematics, March 1996

Seminar, University of Massachusetts, Dept. of Entomology, March 1996

2 Seminars, Utah State University, Dept. of Biology, Jan. 1996

Seminar, Rutgers University, Dept. of Entomology, Sept. 1995

Seminar, Bucknell University, Biology Dept., March 1995

Seminar, University of Arkansas, Dept. of Biological Sciences, Feb. 1995

Seminar, University of Georgia, Depts. of Genetics and Entomology, Jan. 1995

Invited talk, Entomological Society of America meeting, Dallas TX, Dec. 1994 (Symposium title: "Molecular systematics in the service of entomology")

2 Seminars, Field Museum of Natural History, Chicago, IL, Nov. 1994

Plenary lecture, New England Molecular Evolution Meeting, Cambridge, MA, Nov. 1994

Seminar, Hofstra University, Oct. 1994

2 Seminars, University of Florida, Depts. of Zoology and Entomology, Sept. 1994

Seminar, Harvard University, Museum of Comparative Zoology, April 1994

Seminar, Cornell University, Section of Ecology and Systematics, Dec. 1993

Seminar, North Carolina State University, Dept. of Entomology, July 1993

Invited talk, SSE/SSB/ASN meetings, Snowbird UT, June 1993 (Symposium title: "Phylogenetic Studies of Interspecific Interactions")  

 

Contributed Talks and Posters:

Evolution Society meeting, São Paulo, Brazil, June 2015 (coauthor, poster presented by JE Benetti-Longhini)

Evolution Society meeting, Raleigh NC, July 2014 (coauthor, talk presented by IJ Garzón-Orduña) TDAB Andean Butterflies Conference, Urubamba, Peru, Sept. 2008 (2 talks, 1 poster, with Josh Ogawa and Jess Matz)

TLSAMP poster session, MTSU, Aug. 2007 (with Rebecca Guet)

Royal Entomological Society, Bath, U. K. Sept. 2006 (coauthor with LP Brower, K Oberhauser, M Boppré and RI Vane-Wright; paper presented by RIVW)

Lepidopterists’ Society meeting, Gainesville, FL, July 2006 (presented by AD Warren)

Willi Hennig Society meeting, Fagernes Norway, July 2005 (coauthor with K Miller)

Evolution Society meeting, Chico CA June 2003 (coauthor with A Briscoe)

Entomological Society of America National Conference, Ft. Lauderdale FL Nov. 2002 (coauthor with D Murray)

Entomological Society of America National Conference, Montreal, Dec.2000

Lepidopterists’ Society meeting, Wake Forest U., NC, July 2000

Willi Hennig Society meeting, Göttingen, Germany, Sept. 1999

Willi Hennig Society meeting, São Paolo, Brazil, Sept. 1998

Entomological Society of America National Conference, Nashville, TN, Dec. 1997

SSE/SSB/ASN/SNT meetings, Montreal, Quebec, July 1995

SSE/SSB/ASN/SMBE meetings, Athens, GA, June 1994

Entomological Society of America National Conference, Baltimore MD, Dec. 1992

SSE/SSB/ASN meetings, Berkeley, CA, June, 1992

NATO Advanced Science Institute in Molecular Taxonomy, Norwich, U.K. July, 1990  

 

Public Lectures:

Invited speaker, Tennessee Naturalist Program, Cedars of Lebanon State Park, June 2016

Invited speaker, CBAS Research Exchange, Oct. 2015

Invited speaker, Rutherford Recycles, June 2014

Respondent, MTSU Science and Spirituality Forum, Feb. 2013

Invited speaker, Barfield Park Wilderness Station, Oct. 2008

Invited speaker, Kentucky Lepidopterists, Dec. 2006

Invited speaker, Oregon Entomological Society, April 2006

Invited speaker, North American Butterfly Association, Eugene, OR Chapter, Feb. 2006

Invited speaker, Eugene Natural History Society, March 2005

Invited speaker, Washington Butterfly Association, Seattle, Feb. 2002

Keynote Lecture, Oregon Entomological Society, May 2001

Plenary Lecture, Northwest Lepidopterists’ Workshop, Corvallis OR, Oct. 1998

Lecture, TRIAD, Corvallis, OR, Oct. 1998

Lecture, Da Vinci Days, Corvallis, OR, July 1998

Seminar, Entomological Society of Washington, Washington D. C., Oct. 1997

Seminar, New York Butterfly Club, NY, NY, June 1994

Seminar, New York Entomological Society, NY, NY, Feb. 1994

Jordani Lecture, Cornell University, Ithaca, NY, Nov. 1992  

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Awards

Professional Honors:

MTSU Distinguished Research Award, 2012

Entomological Society of America Service Award, 2007

Award for Excellence in Publications, CBAS, MTSU, 2007, 2010

Sigma Xi, 2000

Elected Fellow, Royal Entomological Society, 1999

Elected Fellow, Willi Hennig Society, 1998

Lamont C. Cole Award, Cornell University, 1995

Research/Scholarly Activity

Empirical Systematics and Population Biology:

  • Heliconius systematics, biogeography, coevolution and speciation using molecular and morphological data
  • morphological and molecular cladistics of Ithomiini and Danaini
  • phylogenetic and biogeographical relationships of austral and Andean pronophiline Satyrinae            
  • higher-level systematics of the Nymphalidae using DN...
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Empirical Systematics and Population Biology:

  • Heliconius systematics, biogeography, coevolution and speciation using molecular and morphological data
  • morphological and molecular cladistics of Ithomiini and Danaini
  • phylogenetic and biogeographical relationships of austral and Andean pronophiline Satyrinae            
  • higher-level systematics of the Nymphalidae using DNA sequences           
  • molecular systematics of Hesperiidae, Papilionidae            
  • theory of DNA barcoding
  • molecular systematics of wing-pattern developmental genes in Heliconius butterflies           
  • Danaus plexippus mtDNA population studies            
  • molecular systematics of mimicry in Dioptinae (Notodontidae)            
  • molecular systematics of Hawaiian Drosophila

Systematic Theory:           

  • concept of homology            
  • gene trees vs. species trees
  • time trees and molecular clocks            
  • combining data vs. combining topologies, measures of tree support            
  • philosophy of cladistics            
  • phylogenetic taxonomy and nomenclature            
  • DNA barcoding            
  • parsimony vs. alternative optimality criteria for tree inference
  • "tree-thinking"

Foreign Field Work:            

  • Bahia, Brazil, March 2015
  • Peru, Sept. 2008
  • São Paulo, Brazil March 2004
  • Chile, Jan.-Feb. 1998, Dec. 1998-Jan. 1999
  • Peru, Feb. 1996, Dec. 1996
  • Amazonas and Rondonia, Brazil, Sept.-Oct. 1993
  • Colombia, March 1992
  • Ecuador, Sept. 1990; June 1991
  • Panama, Aug.-Oct. 1990; April 1992
  • French Guiana, May-June 1990; April, 1994
  • Trinidad and Tobago, May 1988
  • Michoacan, Mexico, March 1983; Jan. 1988
  • Puerto Rico, March 1987
  • Dominican Republic, June 1981

Insect Collection Curation:            

  • improvement of MTSU teaching collection, 2015-            
  • Curator of Lepidoptera, Oregon State Arthropod Collection, 1997-2006
  • Visitor/curatorial specialist at:
    • American Museum of Natural History (New York)
    • Smithsonian National Museum of Natural History (Washington, D. C.)
    • Natural History Museum (London)
    • Allyn Museum (Sarasota FL)
    • Naturhistorisches Museum (Wien)
    • Magyar Térmeszet Tudományi Múzeum (Budapest)
    • Musée National de lHistroire Naturelle (Paris)
    • Zoological Museum of the Zoological Institute of the Russian Academy of Sciences (St. Petersburg)
    • Naturalis (Leiden)
    • Staatliches Museum für Naturkunde (Stuttgart)
    • Naturhistoriska Riksmuseet (Stockholm)            
  • Curatorial assistant
    • Cornell University Insect Collection, summer 1989            
    • Graduate Curatorial Fellow, Yale Peabody Museum, 1982-87            
    • Milwaukee Public Museum, summer 1986
    • Harvard Museum of Comparative Zoology, summers 1983, 1984

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In the Media

Torrice, M. 2009. Uncovering Butterflies’ Past. ScienceNOW Daily News, 2 Oct. 2009

Science Channel’s “Through the Wormhole” season 3, “Is there a superior race?” June 2012

Fox News Channel 17 Ant problems on the rise, 31 July 2012

MTSU Magazine 17(2): 14-16 “Imitations in life” featured my research; republished on Entomological Society of America web site, Fall 2012 and Innovations 1:1, F...

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Torrice, M. 2009. Uncovering Butterflies’ Past. ScienceNOW Daily News, 2 Oct. 2009

Science Channel’s “Through the Wormhole” season 3, “Is there a superior race?” June 2012

Fox News Channel 17 Ant problems on the rise, 31 July 2012

MTSU Magazine 17(2): 14-16 “Imitations in life” featured my research; republished on Entomological Society of America web site, Fall 2012 and Innovations 1:1, Fall 2013 (MTSU Basic and Applied Science Magazine)

Wade, L. 2013. The Amazon in 4D. Science 341, 234-235.

Vergano, D. 2014. “Monarch Butterfly's Genes Reveal the Key to Its Long-Distance Migration” (quoted my comments on the research). http://news.nationalgeographic.com/news/2014/10/141001-monarch-migration-genome-science/?utm_source=Twitter&utm_medium=Social&utm_content=link_twngm20141001monarch-migration-genome&utm_campaign=Content&sf4927121=1

Casson, S., and N. Wooton. 2015. “Resting on the Smallest Wings: The Flight of Conservation” Huff Post Generation Change, http://www.huffingtonpost.com/sarah-casson/resting-on-the-smallest-w_b_7107700.html. Posted: 04/21/2015 7:14 pm EDT Updated: 04/21/2015 7:59 pm EDT

Wade, L. 2015. Cradle of life. Science 350, 496-501.

Simon, M. 2016. Twitter nerd-fight reveals a long, bizarre scientific feud. Wired Magazine http://www.wired.com/2016/02/twitter-nerd-fight-reveals-a-long-bizarre-scientific-feud/

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Courses

General Biology 1 (BIOL 1110) Fall 2006; Spring, Fall 2007; Spring, Fall 2008; Spring, Summer, Fall 2009            

General Biology 2 (BIOL 1120) Spring, Summer 2007; Spring 2012; Spring 2013; Spring, Fall 2014; Fall 2015, Spring 2016; Spring 2017

Entomology (BIOL 3040/5040) Fall 2015            

Genetics (BIOL 3250) Fall 2009; Spring, Fall 2010; Spring 2011

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General Biology 1 (BIOL 1110) Fall 2006; Spring, Fall 2007; Spring, Fall 2008; Spring, Summer, Fall 2009            

General Biology 2 (BIOL 1120) Spring, Summer 2007; Spring 2012; Spring 2013; Spring, Fall 2014; Fall 2015, Spring 2016; Spring 2017

Entomology (BIOL 3040/5040) Fall 2015            

Genetics (BIOL 3250) Fall 2009; Spring, Fall 2010; Spring 2011

Evolution (BIOL 3500) Spring, Fall 2010; Spring, Summer, Fall 2011; Spring, Summer, Fall 2012; Spring, Summer 2013; Summer 2014; Spring, Summer 2015, Spring, Summer 2016

Senior Seminar (BIOL 4200) Spring 2017

Evolutionary Psychology (Honors College UH4600) Fall 2008, Spring 2017

Speciation (BIOL 6200) Spring 2008; Fall 2012; Fall 2014

Graduate Seminar BIOL 6650 Fall 2008; Spring 2011; Spring 2012; Fall 2013 “controversies in biology”

Graduate Seminar (BIOL 6660) Fall 2010; Fall 2012 “research tools”

Principles of Systematics (BIOL 6780) Fall 2011, Fall 2013

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